Once had a Porsche 914. Air cooled engine. Drove it across Montana and the Dakotas one winter. One very cold winter.
Not sure the engine ever reached "operating temperature" on that drive.
I've had that happen, too, on a [more] regular car. I drove a Mustang 5.0 from Oklahoma to Oregon, and as I went through eastern Colorado the coolant temperature steadily dropped until it was resting at the bottom of the gauge. I don't recall whether the gas mileage suffered noticeably or not during that phase of the drive.
Sticking a piece of cardboard over a portion of the radiator was a common sight during the winter when I was growing up in rural Ohio. I didn't think our winters were that cold, but maybe late 70s to early 80s vehicles were more susceptible to running cold.
I had a car that developed a stuck-open thermostat and did the cardboard trick to get by until I could replace the faulty part.