They look kind of translucent to me, maybe the first of this kind of slug just had a digestive problem that didn't break down the chloroplasts, and the minimal energy through their bodies made those individuals more successful because they didn't need to eat as often as those who digested theirs. Yada yada other errors among the indegestible-chloroplast population showed further advantages when it's closer to the skin, they outcompeted their peers, etc.