Was the land lower, the seas higher, or some combination, way back when?
Land lower - Alps and Carpathians hadn't formed yet: https://en.wikipedia.org/wiki/Paratethys and its predecessor https://en.wikipedia.org/wiki/Tethys_Ocean
I would wager that the land was lower than the sea back then.
I'm not sure, I would guess both.
Land lower - Alps and Carpathians hadn't formed yet: https://en.wikipedia.org/wiki/Paratethys and its predecessor https://en.wikipedia.org/wiki/Tethys_Ocean