I, personally, would certainly argue that those should be done in small PRs. If you have a working application and want to rewrite in rust, I would expect it to be done by replacing individual components one at a time while making sure relevant tests keep passing. If you're making a new application, I would expect it to be built incrementally by adding parts incrementally.
Yeah, if you ended up doing it as a single massive PR how can you ever be sure that existing functionality was preserved?